李文佳;董彩虹;刘杏忠;李全平;夏金明;梁蕾;

• 1:宜昌山城水都冬虫夏草有限公司
• 2:广东东阳光药业有限公司国家中医药管理局重点研究室
• 3:中国科学院微生物研究所真菌学国家重点实验室

摘要(Abstract)：

本文对近年来冬虫夏草的培植,包括菌种分离培养、寄主饲养、侵染机理、接种方法及子实体发育条件等研究进展进行了综述。本研究组成功实现了冬虫夏草培植产业化,所培植的冬虫夏草与野生冬虫夏草在虫种、菌种、外观形态、显微结构和化学成分等方面一致。最后分析了冬虫夏草培植技术现阶段存在的问题,并对未来进行了展望。

关键词(KeyWords)： 冬虫夏草;接种;子实体发育;菌种分离;寄主饲养

Abstract:

Keywords:

基金项目(Foundation): 国家工信部中药材扶持项目(仿生态培植冬虫夏草产业化生产基地);; 国家自然科学基金(31572179);; 山西省煤基重点科技攻关项目设施食用菌高效碳循环研究与示范(FT2014-03-01)~~

作者(Authors): 李文佳;董彩虹;刘杏忠;李全平;夏金明;梁蕾;

DOI: 10.13346/j.mycosystema.160003

参考文献(References)：

• Chen G,Huang L,Li WJ,Zhan ZP,Xin LB,Ai Z,Qian ZM,Liu GZ,2014.The establishment of NMR characteristic fingerprint of Cordyceps sinensis and its identification study.World Science and Technology/Modernization of Traditional Chinese Medicine and Materia Medica,16(11):2371-2379
• Chen QT,Xiao SR,Shi ZY,1984.Paecilomyces sinensis sp.nov.and its connection with Cordyceps sinensis.Acta Mycologica Sinica,3(1):24-28(in Chinese)
• Chen SJ,Huang TF,Fu SQ,1995.Growth and development of Hepialus gonggaensis.Journal of Chinese Medicinal Materials,18(4):171-173(in Chinese)
• Chen SJ,Yin DH,Li L,Zha X,Shuen JH,Zhama CZ,2000.Resources and distribution of Cordyceps sinensis in Naqu T ibet.Journal of Chinese Medicinal Materials,23(11):673-675(in Chinese)
• Chen SJ,Yin DH,Zhong GY,2003.Biology research of the host Hepialus biruensis eggs of Cordyceps sinensis.Chinese Journal of Traditional Chinese Medicine,28(11):1102-1103(in Chinese)
• Chen YQ,Wang N,Qu LH,Li TH,Zhang WM,2001.Determination of the anamorph of Cordyceps sinensis inferred from the analysis of the ribosomal DNA internal transcribed spacers and 5.8S r DNA.Biochemical Systematics and Ecology,29:597-607(in Chinese)
• Dai RQ,Lan JL,Chen WH,Li XM,1989.Research on Paecilomyces hepiali Chen et Dai.sp.nov.Acta Agriculturae Universitatis Pekinensis,15(2):221-224(in Chinese)
• Dong CH,Li WJ,Li ZZ,Yang WJ,Li TH,Liu XZ,et al.,2016.Cordyceps industry in China:current status,challenges and perspectives-Jinhu declaration for cordyceps industry development.Mycosystema,35(1):1-15(in Chinese)
• Dong CH,Yao YJ,2011.On the reliability of fungal materials used in studies on Ophiocordyceps sinensis.Journal of Industrial Microbiology&Biotechnology,38(8):1027-1035
• Gao L,Li XH,Zhao JQ,Lu JH,Zhu JS,2011.Detection of multiple Ophiocordyceps sinensis mutants in premature stroma of Cordyceps sinensis by Mass-ARRAY SNPMALDI-TOF mass spectrum genotyping.Journal of Peking University(Health Sciences),43(2):259-266(in Chinese)
• Gao ZX,Chen J,Yu H,Zhao ZH,Song HM,1991.Research on artificial cultivation of Cordyceps sinensis-Preliminary report of the host Hepialus of Cordyceps sinensis in Kangding.Journal of Zhejiang Agricultural University,17(1):1-5(in Chinese)
• Ge F,Gui L,Li CR,Fan MZ,2009.Studies on solid-state fementation condition of Hirsutella sinensis anamorph of Cordyceps sinensis.Journal of Biology,3:23-25(in Chinese)
• Guo HC,Gao JQ,Xi QY,Li XH,2003.The research progress of Cordyceps sinensis.Journal of Microbiology,23(1):50-55(in Chinese)
• Guo X,Liu B,Ma SB,Gui MY,2008.The ecological environmental investigation and biological characteristic analysis of Cordyceps sinensis in Yunnan Province.Edible Fungi of China,27(6):8-11(in Chinese)
• Hu X,Zhang YJ,Xiao GH,Zheng P,Xia YL,Zhang XY,Raymond SL,Liu XZ,Wang CS,2013.Genome survey uncovers the secrets of sex and lifestyle in caterpillar fungus.Chinese Science Bulletin,58(23):2846-2854
• Jiang Y,Yao YJ,2002.Names related to Cordyceps sinensis anamorph.Mycotaxon,84:245-254
• Ke CK,2005.Research of Chinese Cordyceps sinensis.China Science and Technology Press,Beijing.1-305(in Chinese)
• Lei W,Peng QY,Zhang GR,Liu X,2015.The latent infection process of Ophiocordyceps sinesis in host Thitarodes larvae.Journal of Environmental Entomology,37(2):387-392(in Chinese)
• Lei W,Shui XR,2014.Study advances on biosystematic of Ophiocordyceps sinesis.Northern Horticulture,2014(14):193-196(in Chinese)
• Li CR,Peng F,Fan MZ,Li ZZ,2004.Studies on liquid and solid culture conditions of Hirsutella sinensis strain RCEF0273,the anamorph of Cordyceps sinensis.Journal of Anhui Agricultural University,31(4):460-465(in Chinese)
• Li L,Yin DH,Chen SJ,2000.The growth and development of Kangding Cordyceps sinensis ascospores.Chinese Journal of Traditional Chinese Medicine,25(8):501-502(in Chinese)
• Li L,Yin DH,Tang GH,Fu SQ,1993.Relationship between the growth and development of Cordyceps sinensis stroma with light.Chinese Journal of Traditional Chinese Medicine,18(2):80-84(in Chinese)
• Li QP,He Y,Liu JM,Xia JM,Li WJ,Liu XZ,2016.Hybrid breeding of high quality of Hepialus sp.,the host of Ophiocordyceps sinensis,and prevention of the host insect reproductive degradation.Mycosystema,35:456-466
• Li QS,Li L,Yin DH,Fu SQ,Huang TF,Luo QM,Fang SN,1991.Biological characteristics of Ophiocordyceps sinensis.Special Wild Economic Animal and Plant Research,1:42-45(in Chinese)
• Li QS,Zeng W,Yin DH,Huang TF,1998.A preliminary study on generations alternation of Cordyceps sinensis fungus.Chinese Journal of Traditional Chinese Medicine,23(4):210-213(in Chinese)
• Li WJ,Wei RS,Xia JM,LüYH,2016.Ophiocordyceps sinensis in the intestines of Hepialus larvae.Mycosystema,35:450-455
• Li ZL,1988.A study of Tolypocladium sinense C.L.Li.sp.nov and cyclosporin production.Acta Mycologica Sinica,7(2):93-98(in Chinese)
• Li ZZ,Huang B,Li CR,Fan MZ,2000.Molecular evidence for anamorph determination of Cordyceps sinensis(Berk.)Sacc.Mycosystema,19(1):60-64(in Chinese)
• Liang ZQ,1994.Cordyceps and artificial culture.Journal of Guizhou Agricultural College,26(Suppl.):1-21(in Chinese)
• Liao ZY,2005.Technical study of the infection of Cordyceps sinesis.Edible Fungi,2005(6):18(in Chinese)
• Liu F,Wu XL,Chen SJ,Yin DH,Zeng W,Zhong GY,2007.Advances in studies on artificial culture of Cordyceps sinensis.Chinese Traditional and Herbal Drugs,38(2):302-305(in Chinese)
• Liu F,Wu XL,Zeng W,Zhang DL,Chen SJ,Yin DH,2008.The influence of integrated pest management technology to the growth and development of the host larvae of Cordyceps sinensis.Chinese Journal of Traditional Chinese Medicine,33(23):2741-2744(in Chinese)
• Liu F,Zeng W,Zhang DL,Yin DH,2013.Studies on death dynamics of artificial cultural host larva of Cordyceps sinensis.Chongqing Journal of Research on Chinese Drugs and Herbs,2013(2):5-6(in Chinese)
• Liu F,Zeng W,Zhang DL,Yin DH,Chen SJ,2009.Comparative studies on the biological character of the Hepialus introduced.Lishizhen Medicine and Materia Medica Research,20(12):3128-3129(in Chinese)
• Liu JL,1990.Anamorph of Cordyceps sinesis and artificial cultivation of the fruiting body.Guizhou Agricultural Science,1990(1):43-48(in Chinese)
• Liu XJ,Guo YL,Yu YX,Zeng W,1989.Separation and identification of Cordyceps fungus agamotype.Journal of Fungi,8(1):35-40(in Chinese)
• Liu ZY,Liang ZQ,Liu AY,Yao YJ,Hyde KD,Yu ZN,2002.Molecular evidence for teleomorph-anamorph connections in Cordyceps based on ITS-5.8S r DNAsequences.Mycological Research,106:1100-1108
• Liu ZY,Yao YJ,Liang ZQ,Liu AY,Pegler DN,Chase MW,2001.Molecular evidence for the anamorph-teleomorph connection in Cordyceps sinensis.Mycological Research,105:827-832
• Lu LX,Liu SM,Shi LG,2002.New advances of study of Cordyceps sinensis.Bulletin of Biology,37(6):4-6(in Chinese)
• Luo YX,2003.The current study situation of Cordyceps sinensis.Journal of Qinghai University(Natural Science Edition),21(2):38-52(in Chinese)
• LüDD,Li ZZ,Wang CS,2008.Advances in molecular pathogenesis and genetic engineering of entomopathogenic fungi.Microbiology China,35(3):443-449(in Chinese)
• Ma SL,Zhang ZH,Xu HF,Liu X,Li YL,2014.Analysis of intestinal flora and infection test of larva,Hepialus lagii.Edible Fungi,2014(3):28-30(in Chinese)
• Mei YX,Yang W,Zhu PX,Peng N,Zhu H,Liang YX,2014.Isolation,characterization,and antitumor activity of a novel heteroglycan from cultured mycelia of Cordyceps sinensis.Planta Medica,80(13):1107-1112
• Qiu Y,Chen YL,Peng C,Wan DG,Shen CH,Yi B,Hou FX,2015.Study on China host insects of Cordyceps sinensis.Lishizhen Medicine and Materia Medica Research,26(3):720-722(in Chinese)
• Shen NY,Zeng L,Zhang XC,Shan Z,Zhou ZR,1983.Isolation of the fungus Cordyceps sinesis.Edible Fungi,1983(5):1-3(in Chinese)
• Sung GH,Hywel-Jones NL,Sung JM,Luangsa-ard JJ,Shrestha B,Spatafora JW,2007.Phylogenetic classification of Cordyceps and the clavicipataceous fungi.Study in Mycology,57:5-59
• Tu YQ,Zhang DL,Zeng W,Chen SJ,2011.Study on biological characteristic of Hepialus xiaojinensis in Sichuan.Chinese Journal of Applied Entomology,48(4):990-996(in Chinese)
• Tu YQ,Zhang DL,Zeng W,Chen SJ,Yin DH,2010.Experimental study of Hepialus larvae infected by Ophicordyceps sinensis.Edible Fungi,2010(3):16-17(in Chinese)
• Tu YQ,Zhu HL,Zhang DL,Chen SJ,2012.Different strains of Cordyceps sinensis on infectivity of host larvae.Edible Fungi of China,31(5):32-34(in Chinese)
• Wang D,2010.Preliminary studies on the bionomics and hand feeding of Hepialus.Master Thesis,Sichuan Agricultural University,Ya′an.30-44(in Chinese)
• Wang GL,Jin HY,Han XP,Shi Y,T ian JG,Lin RC,2008.Quality research and existing problems of medicinal materias Ophiocordyceps sinesis.Chinese Traditional and Herbal Drugs,39(1):115-118(in Chinese)
• Wang HS,2002.The preliminary study on artificial rearing Hepialus spp.Entomological Knowledge,39(2):144-146(in Chinese)
• Wang XL,Yao YJ,2011.Host insect species of Ophiocordyceps sinensis:a review.Zoo Keys,127:43-59
• Wang Z,Ma QL,Qiao ZQ,Ma FQ,2001a.Artificial breeding technology of the host insects Hepialus of Cordyceps sinensis in Gansu.Gansu Agricultural Science and Technology,2001(7):42-43(in Chinese)
• Wang Z,Ma QL,Qiao ZQ,Ma FQ,2001b.Experimental results of artificial innoculaton of Cordyceps sinesis.Gansu Agricultural Science and Technology,2001(7):40-41(in Chinese)
• Wu DT,Meng LZ,Wang LY,Lv GP,Cheong KL,Hu DJ,2014.Chain conformation and immunomodulatory activity of a hyperbranched polysaccharide from Cordyceps sinensis.Carbohydrate Polymers,110:405-414
• Xiang L,Chen SL,Dai Y,Han KH,Wang Q,Chen XP,Zhou ZJ,Ye M,Zhou YJ,2012.Study on biological characteristics of artificial breeding adults of Hepialus xiaojinensis,the larval host of Cordyceps sinensis.World Science and Technology/Modernization of Traditional Chinese Medicine and Materia Medica,14(1):1172-1176(In Chinese)
• Xu HF,2004.Study on biological characteristic of Hepialus lagii in Qinghai.Journal of Sichuan Grassland,2004(106):4-5(in Chinese)
• Xu YD,Xu C,Cui YH,Chao SJ,Du JQ,Liu CC,Zhang LB,2013.The impact of Cordyceps sinensis excavation on grassland vegetation in the Sanjiangyuan region.Research of Environmental Sciences,26(11):1194-1200
• Yang DR,Yang YX,Shen FR,1988.Research on how Cordyceps sinensis fungal infected Hepialus larvae.Chinese Research and Application of Worm Fungus,1988(1):230-234(in Chinese)
• Yang JL,Xiao W,He HX,Zhu HX,Wang SF,Cheng KL,Zhu P,2008.Molecular phylogenetic analysis of Paecilomyces hepiali and Cordyceps sinensis.Acta Phamaceutica Sinica,43(4):421-426(in Chinese)
• Yang YX,Yang DR,Shen FR,Dong DZ,1989.Research on how Cordyceps sinensis fungal infected Hepialus larvae.Zoological Research,10(3):227-231(in Chinese)
• Yin DH,Li L,Huang TF,Luo QM,1994.Ecology of every stage of Hepialus gonggaensis.Special Wild Economic Animal and Plant Research,1994(4):5-7(in Chinese)
• Yin DH,Tang XM,1995.Research progress of artificial culture of Cordyceps sinensis.China Journal of Chinese Materia Medica,20(12):707-709(in Chinese)
• Yin XC,Shen NY,1990.Anamorphs of Cordyceps sinensis-Synnematium sinense Yin&Shen.Journal of High Altitude Biology,9:1-5(in Chinese)
• Yu YX,2004.Research on artificial cultivation of Cordyceps sinensis.Journal of Fungus Research,2(2):42-46(in Chinese)
• Zhang DL,Tu YQ,2015.Ecological research of Hepialus xiaojinensis Tu.Journal of Environmental Insects,37(5):1055-1059(in Chinese)
• Zhang DL,Tu YQ,Liu F,Chen SJ,2014.Study on diet and feeding amount of Hepialus larvae.Journal of Southwest Agriculture,27(6):2732-2734(in Chinese)
• Zhang YJ,Li EW,Wang CS,Li YL,Liu XZ,2012.Ophiocordyceps sinensis,the flag of China:terminology,life strategy and ecology.Mycology,3(1):2-10
• Zhang YJ,Zhang S,Li YL,Ma SL,Wang CS,Xiang MC,Liu X,An ZQ,Xu JP,Liu XZ,2014.Phylogeography and evolution of a fungal-insect association on the T ibetan Plateau.Molecular Ecology,23:5337-5355
• Zhang ZJ,Ye M,Zhou ZJ,Dai Y,Xiang L,2009.Research progress in the breeding of host insects of Cordyceps sinensis.Chinese Traditional and Herbal Drugs,40(Suppl.):85-87(in Chinese)
• Zhao J,Wang N,Chen YQ,Li TH,Qu LH,1999.Molecular identification for the asexual stage of Cordyceps sinensis.Acta Scientiarum Naturalium Universitatis Sunyatseni,38(1):121-123(in Chinese)
• Zhou YJ,Jiang MY,Jiang FX,Liu TM,Zhou ZJ,Zhu TH,2013.Influence of different nitrogen source on Hisurella sinensis growth.Guangdong Agricultural Sciences,40(18):22-24(in Chinese)
• Zhu JS,Guo YL,Yao YS,Zhou YJ,Lu JH,Qi Y,Chen W,Liu XJ,Wu ZM,Zhang L,Yin WT,Zheng TY,Zhang LJ,2007.Coexistence of Hirsutella sinensis and Paecilomyces hepiali DNA during maturation of Cordyceps sinensis and changes in fungal competitive proliferation capability and chemical profiles.Journal of Fungal Research,5(4):214-224(in Chinese)
• Zhu ZG,Liang GH,Liang L,Zhu XY,Li WJ,Xie JJ,Jiang JC,Hua XC,2013.A preparation method of Hirsutella sinensis mature conidia:China,CN103667170A.2014-03-26(in Chinese)
• 陈罡,黄亮,李文佳,詹泽苹,辛立波,艾中,钱正明,刘国柱,2014.冬虫夏草核磁特征指纹图谱建立及鉴别研究.世界科学技术-中医药现代化,16(11):2371-2379
• 陈庆涛,肖生荣,施至用,1984.中国拟青霉新种及其与虫草的关系.真菌学报,3(1):24-28
• 陈仕江,黄天福,付善全,1995.贡嘎蝠蛾生长发育的初步研究.中药材,18(4):171-173
• 陈仕江,尹定华,钟国跃,2003.冬虫夏草寄主比如蝠蛾卵的生物学研究.中国中药杂志,28(11):1102-1103
• 陈仕江,尹定华,李黎,扎西,孙建华,扎玛次仁,2000.西藏那曲地区冬虫夏草资源及分布.中药材,23(11):673-675
• 戴如琴,兰江丽,陈伟华,李晓明,1989.蝙蝠蛾拟青霉新种的研究.北京农业大学学报,15(2):221-224
• 董彩虹,李文佳,李增智,闫文娟,李泰辉,刘杏忠等,2016.我国虫草产业发展现状、问题及展望-虫草产业发展金湖宣言.菌物学报,35(1):1-15
• 高凌,李晓红,赵建晴,陆继红,朱佳石,2011.Mass-ARRAY单核苷酸多态性MALDI-TOF质谱基因分型法检测未成熟冬虫夏草子座中多个冬虫夏草菌突变基因型.北京大学学报(医学版),43(2):259-266
• 高祖紃,陈健,余虹,赵志鸿,宋会鸣,1991.全人工培养冬虫夏草研究-康定虫草寄主蝠蛾研究初报.浙江农业大学学报,17(1):1-5
• 葛飞,桂琳,李春如,樊美珍,2009.冬虫夏草无性型-中国被毛孢固态发酵条件的初步研究.生物学杂志,3:22-25
• 郭宏春,高继全,习欠云,李小红,2003.冬虫夏草研究进展.微生物学杂志,23(1):50-55
• 郭相,刘蓓,马绍宾,桂明英,2008.云南冬虫夏草生态环境调查及生物学特性分析.中国食用菌,27(6):8-11
• 柯传奎,2005.中国冬虫夏草研究.北京:中国科学技术出版社.1-305
• 雷桅,彭青云,张古忍,刘昕,2015.冬虫夏草菌在寄主钩蝠蛾幼虫的潜伏侵染过程研究.环境昆虫学报,37(2):387-392
• 雷桅,税晓容,2014.冬虫夏草生物系统学研究进展.北方园艺,2014(14):193-196
• 李春如,彭凡,樊美珍,李增智,2004.中国被毛孢RCEF0273培养工艺的研究.安徽农业大学学报,31(4):460-465
• 李黎,尹定华,陈仕江,2000.康定冬虫夏草子囊孢子的生长发育.中国中药杂志,25(8):501-502
• 李黎,尹定华,汤国华,付善全,1993.冬虫夏草子座生长发育与光照的关系.中国中药杂志,18(2):80-84
• 李全平,贺媛,刘杰明,夏金明,李文佳,刘杏忠,2016.冬虫夏草寄主昆虫选育及生殖退化研究.菌物学报,35:456-466
• 李泉森,李黎,尹定华,傅善全,黄天福,罗庆明,方苏宁,1991.冬虫夏草的生物学特性.特产研究,1:42-45
• 李泉森,曾伟,尹定华,黄天福,1998.冬虫夏草菌世代交替的初步研究.中国中药杂志,23(4):210-213
• 李文佳,韦瑞升,夏金明,吕延华,2016.蝙蝠蛾幼虫肠道中的冬虫夏草菌体研究.菌物学报,35:450-455
• 李增智,黄勃,李春如,樊美珍,2000.确证冬虫夏草无性型的分子生物学证I.中国被毛孢与冬虫夏草之间的关系.菌物系统,19(1):60-64
• 李兆兰,1988.中国弯颈霉新种及产环孢菌素的研究.真菌学报,7(2):93-98
• 梁宗琦,1994.虫草及其人工培养.贵州农学院丛刊,26(增刊):1-21
• 廖志勇,2005.冬虫夏草的染菌技术研究.食用菌,2005(6):18
• 刘飞,伍晓丽,陈仕江,尹定华,曾纬,钟国跃,2007.冬虫夏草人工培殖的研究概况.中草药,38(2):302-305
• 刘飞,伍晓丽,曾纬,张德利,陈仕江,尹定华,2008.病虫害综合防治技术对冬虫夏草寄主幼虫生长发育的影响.中国中药杂志,33(23):2741-2744
• 刘飞,曾纬,张德利,尹定华,2013.人工饲养冬虫夏草寄主幼虫的死亡动态研究.重庆中草药研究,2013(2):5-6
• 刘飞,曾纬,张德利,尹定华,陈仕江,2009.引种冬虫夏草寄主昆虫的生物学特性比较研究.时珍国医国药,20(12):3128-3129
• 刘杰麟,1990.虫草的无性型及其子实体的人工培养.贵州农业科学,1990(1):43-48
• 刘锡琎,郭英兰,俞永信,曾纬,1989.冬虫夏草无性阶段的分离和鉴定.真菌学报,8(1):35-40
• 陆龙喜,刘淑梅,时连根,2002.冬虫夏草研究的新进展.生物学通报,37(6):4-6
• 罗玉秀,2003.冬虫夏草的研究现状.青海大学学报(自然科学版),21(2):38-52
• 吕丁丁,李增智,王成树,2008.虫生真菌分子致病机理及基因工程改造研究进展.微生物学通报,35(3):443-449
• 马少丽,张宗豪,徐海峰,刘欣,李玉玲,2014.拉脊蝠蛾幼虫肠道菌群分析及侵染试验.食用菌,2014(3):28-30
• 邱乙,程元柳,彭成,万德光,沈才洪,易彬,侯飞霞,国锦琳,2015.中国冬虫夏草寄主昆虫研究.时珍国医国药,26(3):720-722
• 沈南英,曾璐,张显耻,善智,魏世录,周中蓉,1983.冬虫夏草真菌的分离.食用菌,1983(5):1-3
• 涂永勤,张德利,曾纬,陈仕江,2011.四川小金蝠蛾生物学特性研究.应用昆虫学报,48(4):990-996
• 涂永勤,张德利,曾纬,陈仕江,尹定华,2010.蝠蛾属幼虫感染冬虫夏草菌的实验研究.食用菌,2010(3):16-17
• 涂永勤,朱华李,张德利,陈仕江,2012.不同产地冬虫夏草菌对寄主幼虫侵染力研究.中国食用菌,31(5):32-34
• 王栋,2010.蝠蛾生物学性状和人工饲养的初步研究.四川农业大学硕士学位论文,雅安.30-44
• 王钢力,金红宇,韩小萍,石岩,田金改,林瑞超,2008.冬虫夏草药材的质量研究及存在问题.中草药,39(1):115-118
• 王宏生,2002.冬虫夏草蝙蝠蛾人工饲养技术的初步研究.昆虫知识,39(2):144-146
• 王忠,马启龙,乔正强,马福全,2001a.甘肃虫草寄主昆虫蝠蛾的全人工饲养技术.甘肃农业科技,2001(7):42-43
• 王忠,马启龙,乔正强,马福全,2001b.冬虫夏草菌全人工培养感染试验结果.甘肃农业科技,2001(7):40-41
• 向丽,陈士林,代勇,韩开华,王强,陈学平,周祖基,叶萌,周宇爝,2012.冬虫夏草菌寄主小金蝠蛾人工饲养成虫生物学特性.世界科学技术-中医药现代化,14(1):1172-1176
• 徐海峰,2004.青海拉脊蝠蛾的生物学特性观察研究.四川草原,2004(106):4-5
• 徐延达,徐翠,崔永洪,巢世军,杜加强,刘成程,张林波,2013.三江源地区冬虫夏草采挖对草地植被的影响.环境科学研究,26(11):1194-1200
• 杨大荣,杨跃雄,沈发荣,1988.冬虫夏草真菌感染蝠蛾属幼虫的研究.中国虫生真菌的研究与应用,1988(1):230-234
• 杨金玲,肖薇,何惠霞,朱慧新,王淑芳,程克棣,朱平,2008.蝙蝠蛾拟青霉与冬虫夏草关系的分子系统学研究.药学学报,43(4):421-426
• 杨跃雄,杨大荣,沈发荣,董大志,1989.虫草菌感染虫草蝠蛾幼虫的研究.动物学研究,10(3):227-231
• 尹定华,李黎,黄天福,罗庆明,1994.贡嘎蝠蛾各虫态生态学的研究.特产研究,1994(4):5-7
• 尹定华,唐雪梅,1995.冬虫夏草人工培殖研究的进展.中国中药杂志,20(12):707-709
• 印象初,沈南英,1990.冬虫夏草的无性世代-中华束丝孢.高原生物学季刊,9:1-5
• 俞永信,2004.人工培养冬虫夏草研究.菌物研究,2(2):42-46
• 张德利,涂永勤,2015.小金蝠蛾的生态型研究.环境昆虫学报,37(5):1055-1059
• 张德利,涂永勤,刘飞,陈仕江,2014.蝠蛾幼虫奢食性及取食量研究.西南农业学报,27(6):2732-2734
• 张泽锦,叶萌,周祖基,代勇,向丽,2009.冬虫夏草寄主昆虫饲养研究进展.中草药,40(增刊):85-87
• 赵锦,王宁,陈月琴,李泰辉,屈良鹄,1999.冬虫夏草无性型的分子鉴别.中山大学学报自然科学版,38(1):121-123
• 章卫民,李泰辉,陈月琴,屈良鹄,钟韩,徐学平,2002.西藏冬虫夏草无性型的分子生物学研究.微生物学通报,29(3):54-57
• 周宇爝,江明艳,姜福星,刘天美,周祖基,朱天辉,2013.不同氮源对中国被毛孢生长的影响.广东农业科学,40(18):22-24
• 朱佳石,郭英兰,姚艺桑,周妍娇,陆继红,齐莹,谌巍,刘星阶,吴子,张丽,殷蔚荑,郑陶冶,张丽娟,2007.冬虫夏草成熟过程中中国被毛孢和蝙蝠蛾拟青霉DNA共存和竞争增殖力及化学成分指纹谱的变化.菌物研究,5(4):214-224
• 朱志钢,梁关海,梁蕾,朱新燕,李文佳,谢俊杰,蒋均才,华献春,2013.一种制备中国被毛孢成熟分生孢子的方法:China,CN103667170A.2013-09-22